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All 34 Bosses and Their Forms in Monster Train 2

Updated on November 11, 2026 (about 12 hours ago)

Monster Train 2 is a deck-building game set in Heaven, which is split into Rings. You build and upgrade your cards between battles, and every Ring's combat ends with a boss. Bosses are big, tricky enemies with strong effects that can change how the whole fight plays out, and many come in several forms. Soul Savior, a separate mode from the Destiny of the Railforged DLC, has its own set of bosses and a different route through the game.

View

Where

Ring 1

2 items

Astreal the Lost

This Ring 1 boss starts with Sap and gains more when it resolves.
Mode
Standard
Appears In
Ring 1
Forms / Phases
One form
Attack
Base: 8; Covenant 1: 9; Covenant 8: 9
Health
Base: 180; Covenant 1: 360; Covenant 8: 380
Abilities
Base: Relentless. Sap 6.; Covenant 4: Resolve: Gain Sap 1 and restore 30 Health.

Yorael the First-Blessed

This Ring 1 boss builds Rage after slaying a unit.
Mode
Standard
Appears In
Ring 1
Forms / Phases
One form
Attack
Base: 7; Covenant 1: 8; Covenant 4: 8
Health
Base: 125; Covenant 1: 305; Covenant 4: 325
Abilities
Base: Relentless.; Covenant 4: Rage 3. Slay: Gain Rage 1.

Where

Ring 2

2 items

Elebor the Wall

This Ring 2 boss builds Armor during the fight.
Mode
Standard
Appears In
Ring 2
Forms / Phases
One form
Attack
Base: 7; Covenant 1: 8; Covenant 8: 8
Health
Base: 260; Covenant 1: 440; Covenant 8: 460
Abilities
Base: Relentless.; Action: Gain Armor 5.; Covenant 4: Titanskin 2.; Resolve: Gain Damage Shield 1.

Legion

This Ring 2 boss starts with Spikes, so every hit on it hurts the attacker.
Mode
Standard
Appears In
Ring 2
Forms / Phases
One form
Attack
Base: 4; Covenant 1: 5; Covenant 8: 5
Health
Base: 260; Covenant 1: 440; Covenant 8: 460
Abilities
Base: Relentless. Spikes 3.; Covenant 4: Resolve: Gain Spikes 1.

Where

Ring 3

2 items

Arkion, Scion of Savagery

This Ring 3 boss comes in one of three forms, each with a different attack and effect.
Mode
Standard
Appears In
Ring 3
Forms / Phases
Furious Frenzy; Hard Headed; Pyre Hunter
Attack
Base: Furious Frenzy: 7, Hard Headed: 13, Pyre Hunter: 4; Covenant 1: Furious Frenzy: 7, Hard Headed: 13, Pyre Hunter: 4; Covenant 4: Furious Frenzy: 7, Hard Headed: 13, Pyre Hunter: 4; Covenant 8: Furious Frenzy: 7, Hard Headed: 13, Pyre Hunter: 4
Health
Base: Furious Frenzy: 425, Hard Headed: 425, Pyre Hunter: 425; Covenant 1: Furious Frenzy: 625, Hard Headed: 625, Pyre Hunter: 625; Covenant 4: Furious Frenzy: 725, Hard Headed: 725, Pyre Hunter: 725; Covenant 8: Furious Frenzy: 825, Hard Headed: 825, Pyre Hunter: 825
Abilities
Base: Furious Frenzy:; Siphon: Gain Rage 2.; Hard Headed:; Strike: Push the attacked unit to the back.; Pyre Hunter:; Multistrike 1.; Resolve: If you have no units on the floor, deals 5 damage to the Pyre.

Fleshweaver

This Ring 3 boss loses Mass when hit and creates Fleshlings.
Mode
Standard
Appears In
Ring 3
Forms / Phases
Corrupted Mass; Spiked Mass; More Mass
Attack
Base: Corrupted Mass: 6, Spiked Mass: 6, More Mass: 6; Covenant 1: Corrupted Mass: 7, Spiked Mass: 7, More Mass: 7; Covenant 4: Corrupted Mass: 7, Spiked Mass: 7, More Mass: 7; Covenant 8: Corrupted Mass: 7, Spiked Mass: 7, More Mass: 7
Health
Base: Corrupted Mass: 500, Spiked Mass: 500, More Mass: 600; Covenant 1: Corrupted Mass: 680, Spiked Mass: 680, More Mass: 780; Covenant 4: Corrupted Mass: 680, Spiked Mass: 680, More Mass: 780; Covenant 8: Corrupted Mass: 700, Spiked Mass: 700, More Mass: 800
Abilities
Base: Mass 5. Revenge: Create a Fleshling and lose Mass 1.; Corrupted Mass: -; Spiked Mass: -; More Mass: Mass 3.

Where

Ring 4

3 items

Ajax the Unyielding

This Ring 4 boss repeatedly lowers your units’ Attack with Sap.
Mode
Standard
Appears In
Ring 4
Forms / Phases
One form
Attack
Base: 9; Covenant 1: 12; Covenant 4: 12; Covenant 8: 12
Health
Base: 550; Covenant 1: 800; Covenant 4: 900; Covenant 8: 1000
Abilities
Base: Relentless. Action: Apply Sap 3 to your units.; Covenant 4: Strike: Apply Sap 3.

Korin the Judge

This Ring 4 boss pulls surviving enemies to her floor and retreats.
Mode
Standard
Appears In
Ring 4
Forms / Phases
One form
Attack
Base: 8; Covenant 1: 11; Covenant 4: 11; Covenant 8: 11
Health
Base: 600; Covenant 1: 850; Covenant 4: 950; Covenant 8: 1050
Abilities
Base: Relentless. Summon: Descend all friendly units to this floor and Retreat.; Covenant 4: Summon: Apply Multistrike 1 to all enemy units.

Qel the Corruptor

This Ring 4 boss applies Corruption when units cast spells.
Mode
Standard
Appears In
Ring 4
Forms / Phases
One form
Attack
Base: 12; Covenant 1: 15; Covenant 4: 15; Covenant 8: 15
Health
Base: 700; Covenant 1: 950; Covenant 4: 1050; Covenant 8: 1150
Abilities
Base: Relentless.; Incant: Apply Corruption 3 to your units.; Covenant 4: Summon: Apply Corruption 5 to all your units.

Where

Ring 5

2 items

Quoto the Absolute

This Ring 5 boss starts with heavy Titanskin and loses stacks when hit.
Mode
Standard
Appears In
Ring 5
Forms / Phases
One form
Attack
Base: 16; Covenant 1: 19; Covenant 4: 19; Covenant 8: 19
Health
Base: 900; Covenant 1: 1150; Covenant 4: 1250; Covenant 8: 1350
Abilities
Base: Relentless.; Titanskin 40.; Revenge: Lose Titanskin 1.; Covenant 4: Resolve: Gain Titanskin 5.

The Undying One

This Ring 5 boss uses Reanimate to prolong its fight.
Mode
Standard
Appears In
Ring 5
Forms / Phases
One form
Attack
Base: 17; Covenant 1: 20; Covenant 4: 20; Covenant 8: 20
Health
Base: 500; Covenant 1: 750; Covenant 4: 850; Covenant 8: 950
Abilities
Base: Relentless. Reanimate 8.; Covenant 4: Resolve: Gain Reanimate 6.

Where

Ring 6

3 items

Cael, Lord of the Cherubs

This Ring 6 boss summons Cherubs that grant effects when eaten.
Mode
Standard
Appears In
Ring 6
Forms / Phases
Holy Feed; Nourishing Feed; Vampiric Feed
Attack
Base: 22; Covenant 1: 22; Covenant 4: 22; Covenant 8: 22
Health
Base: 1800; Covenant 1: 2500; Covenant 4: 2800; Covenant 8: 3100
Abilities
Base: Holy Feed:; Resolve: Grant Damage Shield 1 to the front non-boss enemy unit.; Nourishing Feed:; Resolve: Grant +35 Health and Spikes 4 to the front non-boss enemy unit.; Vampiric Feed:; Resolve: Grant +7 Attack and Lifesteal 1 to the front non-boss enemy unit.

Hallow

This Ring 6 boss shares a fight with Stryx and supports her with changing effects.
Mode
Standard
Appears In
Ring 6
Forms / Phases
Bloody Harvest; Doombringers; Inevitability
Attack
Base: Bloody Harvest: 11, Doombringers: 11, Inevitability: 11; Covenant 1: Bloody Harvest: 14, Doombringers: 14, Inevitability: 14; Covenant 4: Bloody Harvest: 14, Doombringers: 14, Inevitability: 14; Covenant 8: Bloody Harvest: 14, Doombringers: 14, Inevitability: 14
Health
Base: Bloody Harvest: 1250, Doombringers: 1250, Inevitability: 1250; Covenant 1: Bloody Harvest: 1500, Doombringers: 1500, Inevitability: 1500; Covenant 4: Bloody Harvest: 1600, Doombringers: 1600, Inevitability: 1600; Covenant 8: Bloody Harvest: 1700, Doombringers: 1700, Inevitability: 1700
Abilities
Base: Titanskin 10. Extinguish: Stryx gains Titanskin 10. Relentless begins next turn.; Bloody Harvest: Siphon: Stryx gains +50 Health.; Doombringers: Incant: Stryx gains +30 Health.; Inevitability: Resolve: Stryx gains +60 Health.

Stryx

This Ring 6 boss shares a fight with Hallow and keeps adding Attack to Hallow as the fight goes on.
Mode
Standard
Appears In
Ring 6
Forms / Phases
Bloody Harvest; Doombringers; Inevitability
Attack
Base: Bloody Harvest: 13, Doombringers: 13, Inevitability: 13; Covenant 1: Bloody Harvest: 16, Doombringers: 16, Inevitability: 16; Covenant 4: Bloody Harvest: 16, Doombringers: 16, Inevitability: 16; Covenant 8: Bloody Harvest: 16, Doombringers: 16, Inevitability: 16
Health
Base: Bloody Harvest: 1100, Doombringers: 1100, Inevitability: 1100; Covenant 1: Bloody Harvest: 1350, Doombringers: 1350, Inevitability: 1350; Covenant 4: Bloody Harvest: 1450, Doombringers: 1450, Inevitability: 1450; Covenant 8: Bloody Harvest: 1550, Doombringers: 1550, Inevitability: 1550
Abilities
Base: Multistrike 1. Live to Fight. Extinguish: Hallow gains Multistrike 1. Relentless begins next turn.; Bloody Harvest: Revenge: Hallow gains +1 Attack.; Doombringers: Slay: Hallow gains +3 Attack.; Inevitability: Loop: Hallow gains +8 Attack.

Where

Ring 7

2 items

Athane, Last to Fall

This Ring 7 boss summons a Chosen Prayerstone and grants Spikes to enemy units.
Mode
Standard
Appears In
Ring 7
Forms / Phases
One form
Attack
Base: 12; Covenant 1: 17; Covenant 4: 17; Covenant 8: 17
Health
Base: 1100; Covenant 1: 1475; Covenant 4: 1725; Covenant 8: 2125
Abilities
Base: Relentless.; Incant: Apply Spikes 1 to enemy units.; Summon Ascension: Spawn a Chosen Prayerstone.

Rax the Overblessed

This Ring 7 boss attacks twice and gains Rage, Regen and Spell Shield each time it resolves.
Mode
Standard
Appears In
Ring 7
Forms / Phases
One form
Attack
Base: 9; Covenant 1: 14; Covenant 4: 14; Covenant 8: 14
Health
Base: 1100; Covenant 1: 1475; Covenant 4: 1725; Covenant 8: 2125
Abilities
Base: Multistrike 1.; Rage 8; Regen 20.; Spell Shield 2.; Resolve: Gain Rage 8, Regen 20, and Spell Shield 2.; Covenant 4: Resolve: Gain Multistrike 1.

Where

Ring 8

1 items

Seraph Aeternus

This Ring 8 boss appears in one of three Titan-themed forms.
Mode
Standard
Appears In
Ring 8
Forms / Phases
The Dominant; The Entropic; The Savage
Attack
The Dominant: Base: 20; Covenant 1: 20; Covenant 4: 20; Covenant 8: 20; The Entropic: Base: 35; Covenant 1: 35; Covenant 4: 35; Covenant 8: 35; The Savage: Base: 30; Covenant 1: 30; Covenant 4: 30; Covenant 8: 30
Health
The Dominant: Base: 2800; Covenant 1: 3800; Covenant 4: 5300; Covenant 8: 6300; The Entropic: Base: 3800; Covenant 1: 4800; Covenant 4: 6300; Covenant 8: 7300; The Savage: Base: 3000; Covenant 1: 4000; Covenant 4: 5500; Covenant 8: 6500
Abilities
The Dominant — Base: Relentless.; Strike: Double Corruption on the attacked unit.; Action: Apply Corruption 2 to your units.; The Entropic — Base: Relentless.; Resolve: Add a Curse of Entropy card to your deck.; Enchant: Grant your units Sap 5.; The Savage — Base: Relentless.; Enchant: Grant enemy units Rage 3.

Where

Ring 9

3 items

Dominion

This Ring 9 Titan stays by the Pyre and adds Corruption to your front units.
Mode
Standard
Appears In
Ring 9
Forms / Phases
One form
Attack
Base: 3; Covenant 4: 3
Health
Base: 1555; Covenant 4: 2055
Abilities
Base: Fate Link. Immobile. Pyre Bound. Spell Shield 4.; Resolve: Add Pyre Light to your hand. Apply Corruption 5 to your front unit on all floors.

Entropy

This Ring 9 Titan never moves and adds a Curse of Entropy to your deck each time it resolves.
Mode
Standard
Appears In
Ring 9
Forms / Phases
One form
Attack
Base: 15; Covenant 4: 20
Health
Base: 3500; Covenant 4: 4500
Abilities
Base: Fate Link.; Immobile.; Resolve: Add a Curse of Entropy to your deck.

Savagery

This Ring 9 Titan tramples through your units, builds Rage and hits the Pyre directly when its floor is empty.
Mode
Standard
Appears In
Ring 9
Forms / Phases
One form
Attack
Base: 10; Covenant 4: 15
Health
Base: 2750; Covenant 4: 3250
Abilities
Base: Fate Link.; Live to Fight.; Trample.; Resolve: Gain Rage 2. If you have no units on the floor, deals 5 damage to the Pyre.; Loop: Gain Multistrike 1.

Where

Soul Savior: First Battle

1 items

Astrael the First Reborn

ATAstrael the First Reborn
The opening Soul Savior boss starts with Sap and Infested, and its stats rise with the mode difficulty.
Mode
Soul Savior
Appears In
Soul Savior: First Battle
Forms / Phases
One form
Attack
Bloom: 8; Tangle: 12; Overgrowth: 16
Health
Bloom: 230; Tangle: 345; Overgrowth: 460
Abilities
Relentless; starts with Sap 6 and Infested 4.

Where

Soul Savior: Maera's Region

3 items

Athane the Fallen

ATAthane the Fallen
This first-combat boss in Maera’s Region applies Rage to enemies and summons an Ossivane Sentinel.
Mode
Soul Savior
Appears In
Soul Savior: Maera's Region
Forms / Phases
One form
Attack
Bloom: 12; Tangle: 18; Overgrowth: 24
Health
Bloom: 400; Tangle: 600; Overgrowth: 800
Abilities
Relentless; Incant: apply Rage 1 to enemy units; Ascension: spawn an Ossivane Sentinel; Strike: Retreat.

Korin the Judged

KTKorin the Judged
This first-combat boss in Maera’s Region pulls surviving friendly units to its floor before retreating.
Mode
Soul Savior
Appears In
Soul Savior: Maera's Region
Forms / Phases
One form
Attack
Bloom: 12; Tangle: 18; Overgrowth: 24
Health
Bloom: 400; Tangle: 600; Overgrowth: 800
Abilities
Relentless; Summon: Descend all friendly units to this floor and Retreat.

Maera the Dutiful

The flying boss of Maera’s Region has three forms that grant Armor, Rage, or Burst to enemies.
Mode
Soul Savior
Appears In
Soul Savior: Maera's Region
Forms / Phases
Eldest Scion; Sibling Hierarchy; Stern Sister
Attack
Eldest Scion: 12; Sibling Hierarchy: 15; Stern Sister: 16
Health
Eldest Scion: 1000; Sibling Hierarchy: 900; Stern Sister: 800
Abilities
Eldest Scion: Resolve grants enemy units Armor equal to their Attack. Sibling Hierarchy: Shift applies Rage 3/4/5 by difficulty. Stern Sister: Resolve applies Burst 1 to Maera and enemy units.

Where

Soul Savior: Thaddeus's Region

3 items

Elebor the Unstoppable

ETElebor the Unstoppable
This is one of the two first-combat boss choices in Thaddeus’s Region.
Mode
Soul Savior
Appears In
Soul Savior: Thaddeus's Region
Forms / Phases
-
Abilities
-

Quoto the Destroyer

QTQuoto the Destroyer
This is one of the two first-combat boss choices in Thaddeus’s Region.
Mode
Soul Savior
Appears In
Soul Savior: Thaddeus's Region
Forms / Phases
-
Abilities
-

Thaddeus the Indulgent

The flying boss of Thaddeus’s Region has three forms built around Capacity loss or Gorge.
Mode
Soul Savior
Appears In
Soul Savior: Thaddeus's Region
Forms / Phases
Train Chomper; Thick Skinned; Spoiled
Abilities
Train Chomper: reduces floor Capacity by 1 and triggers Gorge on Resolve. Thick Skinned: starts with Titanskin 3 and gains Titanskin 3 on Gorge. Spoiled: triggers Gorge on Slay.

Where

Soul Savior: Tivi's Region

3 items

Phalanx

PPhalanx
This is one of the two first-combat boss choices in Tivi’s Region.
Mode
Soul Savior
Appears In
Soul Savior: Tivi's Region
Forms / Phases
-
Abilities
-

The Undying Spirit

TUThe Undying Spirit
This is one of the two first-combat boss choices in Tivi’s Region.
Mode
Soul Savior
Appears In
Soul Savior: Tivi's Region
Forms / Phases
-
Abilities
-

Tivi the Unruly

The flying boss of Tivi’s Region has three forms that copy enemies, advance units, or add Scourge cards.
Mode
Soul Savior
Appears In
Soul Savior: Tivi's Region
Forms / Phases
Duplicitous; Mischevious Child; Prankster
Abilities
Duplicitous: creates a copy of the front enemy unit on the floor at turn end, or a Jeermask if no enemies are there. Mischevious Child: has Sniper and Advances the attacked unit on Strike. Prankster: adds a Tivi’s Scourge card to your hand on Siphon.

Where

Soul Savior: Lylith's Region

3 items

Ajax the Deathbringer

ATAjax the Deathbringer
This is one of the two first-combat boss choices in Lylith’s Region.
Mode
Soul Savior
Appears In
Soul Savior: Lylith's Region
Forms / Phases
One form
Abilities
-

Lylith the Spurned

The flying boss of Lylith’s Region has three forms that use Dualism, Witherbloom, or Armor.
Mode
Soul Savior
Appears In
Soul Savior: Lylith's Region
Forms / Phases
Plaguebringer; Energy Vampire; Inoculation
Abilities
Plaguebringer: gains Dualism at 50% health. Energy Vampire: applies Witherbloom to your units on Action. Inoculation: gains Armor equal to 3 times her Malaise on Strike.

Qel the Malaiser

QTQel the Malaiser
This is one of the two first-combat boss choices in Lylith’s Region.
Mode
Soul Savior
Appears In
Soul Savior: Lylith's Region
Forms / Phases
One form
Abilities
-

Where

Soul Savior: Final Battle

1 items

The Lifemother

The final Soul Savior encounter has three forms, each with a different effect on the train.
Mode
Soul Savior
Appears In
Soul Savior: Final Battle
Forms / Phases
The Corpseflower; The Swarmhost; The Undying Bloom
Abilities
The Corpseflower: applies Malaise 12 on Strike and spreads Debuff effects on Hellborne Harvest. The Swarmhost: gains Infested 2 on Harvest. The Undying Bloom: applies Witherbloom 10 to the front friendly unit and Reanimate 2 to the front enemy unit at turn end.

Standard Run Bosses by Ring

In standard runs, the kind of boss changes as you move up the Rings. Rings 1, 2, 4, 5 and 7 hold minor bosses, which are regular enemies that appear on the final wave of their fight with the Relentless effect. Rings 3, 6 and 8 hold the major bosses, and Ring 9 holds the Titans.

Ring 3 is home to Arkion and Fleshweaver, Ring 6 has Cael and the Stryx and Hallow pair, and Ring 8 brings the Seraph Aeternus forms. Ring 9 sits outside the regular climb. It only opens once you have collected every Titan Essence, and fighting it after Ring 8 is an optional extra battle that gives a higher score and extra unlocks.

Standard boss stats also depend on your Covenant Rank. Health generally climbs as the rank goes up, and Attack often rises too. That means the same boss can hit harder at a higher rank.

Flying Bosses and How They Move

Most of the major bosses on Rings 3, 6 and 8 are flying bosses, so they don't stay on one floor. Stryx is also a major boss, but its movement is different.

A flying boss doesn't wait for the final wave. It appears on turn 1 and moves to a different floor after every turn, and it only becomes a regular, non-flying boss with the Relentless effect once the final wave starts.

On turn 1, it appears on the highest floor that already has your units during deployment. If you've placed units on every floor, it starts on the top floor. Units summoned at the start of turn 1 by certain artifacts don't count toward this.

After each turn, the boss prefers to fly to a floor that has your units. When all three floors stay occupied, its route settles into a repeating loop that always moves either up the floors or down them. If only one or two floors have your units, it keeps returning to them.

That makes your deployment a real choice. If you want to face the boss on your main unit floor as often as possible, occupy only one or two floors. If you want to face it there as rarely as possible, as you would against Seraph Aeternus-The Dominant, fill all three floors.

Once the Relentless phase begins, some effects switch off. Arkion's triggers stop working, though his Furious Frenzy form keeps any Rage it has built, and Seraph Aeternus-The Dominant stops applying Corruption to all your units with its action while still doubling Corruption on the unit it attacks. Most other effects keep working.

Boss Effects to Plan For

Boss effects fall into a few broad groups, and knowing which one you're facing shapes how you prepare.

Some bosses punish you for hitting them. Legion starts with Spikes, so every hit on it hurts the attacker. Quoto the Absolute begins with heavy Titanskin that drops each time it's struck, and Fleshweaver loses Mass when hit and creates Fleshlings.

Others wear down your units over time. Ajax the Unyielding keeps applying Sap, which steadily lowers the Attack of your units, and Qel the Corruptor adds Corruption to your units whenever you play a spell on its floor. Dominion, one of the Titans, adds Corruption to your front unit on every floor.

Still others get stronger as the fight goes on. Elebor the Wall piles on Armor during the fight, and Yorael the First-Blessed builds Rage each time it slays a unit. The Undying One uses Reanimate to stretch its fight, and Rax the Overblessed gains Rage, Regen and Spell Shield every time it resolves.

Some bosses bring their own helpers. Cael summons Cherubs that grant effects when eaten, and Athane, Last to Fall summons a Chosen Prayerstone.

Others reach into your deck. Entropy adds a Curse of Entropy card to your deck each time it resolves, and the Entropic form of Seraph Aeternus does the same.

Bosses That Matter Most

Some bosses stand out for their numbers, and others for how their fights unfold. The Entropic form of Seraph Aeternus on Ring 8 has the highest Attack and Health of any standard boss. The other forms on Ring 8 play very differently, so it helps to know which one you're facing.

The Titans on Ring 9 each fight in their own way. Dominion stays put by the Pyre, and Entropy never moves while having both the most Attack and the most Health of the Titans. Savagery tramples through your units and can damage the Pyre directly when its floor is empty.

Stryx and Hallow share one fight on Ring 6, and each one makes the other stronger. Hallow's effects keep adding Health to Stryx, while Stryx's effects keep adding Attack to Hallow, so the pair can grow stronger as the fight goes on.

How Soul Savior Bosses Work

Soul Savior runs start with a first battle against Astrael, the First Reborn. After that, the regions can be cleared in any order, and each region ends with a battle against its boss. The run finishes with a final battle once the regions are done.

Soul Savior has its own difficulty levels, labeled Bloom, Tangle and Overgrowth, and boss numbers rise with each one. Region bosses also grow stronger as you complete more regions, so the ones you face later in a run tend to be tougher.

Some Soul Savior bosses share a name with a standard boss, such as Korin, Athane, Qel and Quoto, but they are separate enemies with their own stats and abilities. Treat each one as its own fight, even if the name looks familiar.

Soul Savior Region Bosses

Each region offers a choice of first-combat boss, and its major boss waits at the end. In Maera's Region you can meet Athane the Fallen or Korin the Judged. In Thaddeus's Region it's Elebor the Unstoppable or Quoto the Destroyer, in Tivi's Region it's Phalanx or The Undying Spirit, and in Lylith's Region it's Ajax the Deathbringer or Qel the Malaiser.

Every region's major boss is a flying boss, and each comes in forms built around a theme. Maera the Dutiful's forms hand out Armor, Rage or Burst to the enemy units around her, and Thaddeus the Indulgent's forms work through floor Capacity and Gorge. Tivi the Unruly's forms copy enemy units, advance your units or add Scourge cards to your hand, and Lylith the Spurned's forms use Dualism, Witherbloom or Armor.

FAQ

Q.

Do you still get a Boss Artifact for beating a Soul Savior boss?

No. In Soul Savior, a boss kill gives you an extra Soul instead of the Boss Artifact you'd receive in a standard run.

Q.

What happens after you beat a Soul Savior region boss?

You receive a negative mutator, and one central map node in every other region is permanently upgraded to a stronger "+" version. Once you've beaten the bosses of all but one region, every central node in the last region has been upgraded.

Q.

Can you move flying bosses with Advance, Retreat or Rooted?

Not with those effects. Flying bosses are immune to Advance, Retreat and Rooted, but Ascend and Descend can still move them. Ascend can't push one into the Pyre room before its Relentless phase, and a forced move can flip its flight pattern.

Q.

How long does a flying boss keep flying before the Relentless phase?

In the Arkion and Fleshweaver fights, the boss gets 5 flying turns before the Relentless phase. In the Cael, Stryx and Hallow, and Seraph Aeternus fights, it gets 7.

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